Multi-tuberous

	† Multi-tuberous
	; Skull Ptilodus
Scientific classification
Domain:	Eukaryotes
Kingdom:	Animals
Kingdom :	Eumetazoi
No rank :	Bilateral symmetrical
No rank :	Secondary
Type of:	Chordate
Subtype :	Vertebrates
Infratype :	Maxillary
Overclass :	Tetrapods
Grade:	Mammals
Subclass :	† Alloterias
Squad:	† Multi-tuberous
International scientific name
	Multituberculata Cope , 1884
Suborders
	† Cimolodonta McKenna, 1975; † Plagiaulacida Ameghino, 1889;
23.03	Neogene
66.0	Paleogen
199.6	Yura
251	Triassic
359.2	Carbon
416	Devonian
443.7	Silur
488.3	Ordovician
542	Cambrian
4570	Precambrian

Many-bellied , or multi- tuberculates ^[1] ( lat. Multituberculata ) , - a squad of extinct mammals from the subclass of allotheria (Allotheria). The name comes from the shape of molars , on which numerous tubercles were located. Outwardly similar to rodents . They were distributed only in the Northern Hemisphere on the mainland Laurasia .

They arose at the end of the Triassic period and became extinct by the Oligocene (208.5–33.9 million years ago), having existed for almost 175 million years ^[2] .

More than 200 species are known, ranging in size from mouse to beaver. These species occupied various ecological niches: from inhabitants of burrows to protein-like inhabitants of tree crowns and jerboa-shaped bipedal jumpers ^[3] ^[4] .

Systematics

Many-tuberous ones are usually placed in the mammalian tree as a crown group (or crown group) outside of each of the two main groups of living mammals: therias (including placental and marsupials) and single-pass ^[5] , but closer to the thorium than to single-pass ^[6] ^{[ 7]} .

Mammalia

Australosphenida

Boreosphenida

Fruitafossor

Eutriconodonta

Allotheria

Tinodon

Haramiyavia

Arboroharamiya

Multituberculata

	Plagiaulacida

	Cimolodonta

Symmetrodonta

	Dryolestes

	Henkelotherium

	Amphitherium

	Peramus

Vincelestes

Nanolestes

Aegialodon

Theria

	Metatheria

	Eutheria

Classification

The list is based on a combination of data from the Mikko Phylogenetic Archive ^[8] and Paleofile.com. ^[9]

Suborder † Plagiaulacida Simpson 1925

Genus? † Argillomys Cifelli, Gordon & Lipka 2013
- View † Argillomys marylandensis Cifelli, Gordon & Lipka 2013
Genus? † Janumys Eaton & Cifelli 2001
- View † Janumys erebos Eaton & Cifelli 2001
Superfamily † Allodontoidea Marsh 1889
- Genus †? Glirodon Engelmann & Callison, 2001
  - Species † G. grandis Engelmann & Callison, 2001
- Family † Arginbaataridae Hahn & Hahn, 1983
  - Genus Arginbaatar Trofimov, 1980
    - Species † A. dmitrievae Trofimov, 1980
- Family † Zofiabaataridae Bakker, 1992
  - Genus † Zofiabaatar Bakker & Carpenter, 1990
    - View † Z. pulcher Bakker & Carpenter, 1990
- Family † Allodontidae Marsh, 1889
  - Genus Passumys Cifelli, Davis & Sames 2014
    - View † Passumys angelli Cifelli, Davis & Sames 2014
  - Genus † Ctenacodon Marsh, 1879
    - Species † C. serratus Marsh, 1879
    - Species † C. nanus Marsh, 1881
    - View † C. laticeps (Marsh, 1881) [ Allodon laticeps Marsh 1881 ]
    - View † C. scindens Simpson, 1928
  - Genus † Psalodon Simpson, 1926
    - Species † P. potens (Marsh, 1887) [ Ctenacodon potens Marsh 1887 ]
    - View † P. fortis (Marsh, 1887) Simpson 1929 [ Allodon fortis Marsh 1887 ]
    - View † P. marshi Simpson, 1929
Superfamily † Paulchoffatioidea Hahn 1969 sensu Hahn & Hahn 2003
- Genus? † Mojo Hahn, LePage & Wouters 1987
  - View † Mojo usuratus Hahn, LePage & Wouters 1987
- Genus? † Rugosodon Yuan et al. , 2013
  - Species † Rugosodon eurasiaticus Yuan et al. , 2013
- Family † Pinheirodontidae Hahn & Hahn, 1999
  - Genus Bernardodon Hahn & Hahn, 1999
    - Species † B. atlanticus Hahn & Hahn, 1999
    - Species † B. sp. Hahn & Hahn, 1999
  - Genus † Cantalera Badiola, Canudo & Cuenca-Bescos, 2008
    - View † Cantalera abadi Badiola, Canudo & Cuenca-Bescos, 2008
  - Genus † Ecprepaulax Hahn & Hahn, 1999
    - Species † E. anomala Hahn & Hahn, 1999
  - Genus † Gerhardodon Kielan-Jaworowska & Ensom, 1992
    - Species † G. purbeckensis Kielan-Jaworowska & Ensom, 1992
  - Genus † Iberodon Hahn & Hahn, 1999
    - Species † I. quadrituberculatus Hahn & Hahn, 1999
  - Genus † Lavocatia Canudo & Cuenca-Bescós, 1996
    - Species † L. alfambrensis Canudo & Cuenca-Bescós, 1996
  - Genus † Pinheirodon Hahn & Hahn, 1999
    - Species † P. pygmaeus Hahn & Hahn, 1999
    - Species † P. vastus Hahn & Hahn, 1999
- Family † Paulchoffatiidae Hahn, 1969
  - Genus? † Galveodon Hahn & Hahn, 1992
    - Species † G. nannothus Hahn & Hahn, 1992
  - Genus † † Sunnyodon Kielan-Jaworowska & Ensom, 1992
    - Species † S. notleyi Kielan-Jaworowska & Ensom, 1992
  - Subfamily † Paulchoffatiinae Hahn, 1971
    - Genus † Paulchoffatia Kühne, 1961
      - View † P. delgador Kühne, 1961
    - Genus † Pseudobolodon Hahn, 1977
      - Species † P. oreas Hahn, 1977
      - Species † P. krebsi Hahn & Hahn, 1994
    - Genus † Henkelodon Hahn, 1987
      - Species † H. naias Hahn, 1987
    - Genus † Guimarotodon Hahn, 1969
      - Species † G. leiriensis Hahn, 1969
    - Genus † Meketibolodon (Hahn, 1978) Hahn, 1993
      - Species † M. robustus (Hahn, 1978) Hahn, 1993 [ Pseudobolodon robusutus Hahn 1978 ]
    - Genus † Plesiochoffatia Hahn & Hahn, 1999 [ Parachoffatia Hahn & Hahn 1998 non Mangold 1970 ]
      - Species † P. thoas (Hahn & Hahn, 1998) Hahn & Hahn 1999 [ Parachoffatia thoa Hahn & Hahn 1998 ]
      - Species † P. peparethos (Hahn & Hahn, 1998) Hahn & Hahn 1999 [ Parachoffatia peparethos Hahn & Hahn 1998 ]
      - Species † P. staphylos (Hahn & Hahn, 1998) Hahn & Hahn 1999 [ Parachoffatia staphylos Hahn & Hahn 1998 ]
    - Genus † Xenachoffatia Hahn & Hahn, 1998
      - View † X. oinopion Hahn & Hahn, 1998
    - Genus † Bathmochoffatia Hahn & Hahn, 1998
      - Species † B. hapax Hahn & Hahn, 1998
    - Genus † Kielanodon Hahn, 1987
      - View † K. hopsoni Hahn, 1987
    - Genus † Meketichoffatia Hahn, 1993
      - Species † M. krausei Hahn, 1993
    - Genus † Renatodon Hahn, 2001
      - View † Renatodon amalthea Hahn, 2001
  - Subfamily † Kuehneodontinae Hahn, 1971
    - Genus † Kuehneodon Hahn, 1969
      - View † K. dietrichi Hahn, 1969
      - Species † K. barcasensis Hahn & Hahn, 2001
      - Species † K. dryas Hahn, 1977
      - Species † K. guimarotensis Hahn, 1969
      - View † K. hahni Antunes, 1988
      - View † K. simpsoni Hahn, 1969
      - Species † K. uniradiculatus Hahn, 1978
Superfamily † Plagiaulacoidea Ameghino, 1894
- Family † Plagiaulacidae Gill, 1872 sensu Kielan-Jaworowska & Hurum, 2001 [Bolodontidae Osborn 1887 ]
  - Genus? † Morrisonodon Hahn & Hahn, 2004
    - Species † Morrisonodon brentbaatar (Bakker, 1998) Hahn & Hahn, 2004 [ Ctenacodon brentbaatar Bakker, 1998 ]
  - Genus † Plagiaulax Falconer, 1857
    - Species † P. becklesii Falconer, 1857
    - Species † P. dawsoni Woodward, 1891 [ Plioprion dawsoni Woodward, 1891 ; Loxaulax dawsoni (Woodward, 1891) Sloan, 1979 ]
  - Genus † Bolodon Owen, 1871 [ Plioprion Cope, 1884 ]
    - Species † B. crassidens Owen, 1871
    - Species † B. falconeri Owen, 1871 [ Pligiaulax falconeri Owen, 1871 ; Plioprion falconeri (Owen, 1871) ]
    - Species † B. hydei Cifelli, Davis & Sames, 2014
    - Species † B. minor Falconer, 1857 [ Pligiaulax minor Falconer, 1857 ; Plioprion minor (Falconer, 1857) ]
    - Species † B. osborni Simpson, 1928 [ Plioprion osborni (Simpson, 1928) ; Ctenacodon osborni Simpson, 1928 ]
    - Species? † B. elongatus Simpson, 1928
Family † Eobaataridae Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
- Genus † Eobaatar Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
  - Species † E. clemensi Sweetman, 2009
  - Species † E. hispanicus Hahn & Hahn, 1992
  - Species † E. magnus Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
  - Species † E. minor Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
  - Species † E. pajaronensis Hahn & Hahn, 2001
- Genus † Hakusanobaatar Kusuhashi et al. , 2008
  - Species † H. matsuoi Kusuhashi et al. , 2008
- Genus † Heishanobaatar Kusuhashi et al. , 2010
  - Species † H. triangulus Kusuhashi et al. , 2010
- Genus † Iberica Badiola et al. , 2011
  - Species † Iberica hahni Badiola et al. , 2011
- Genus † Liaobaatar Kusuhashi et al., 2009
  - Species † L. changi Kusuhashi et al. , 2009
- Genus † Loxaulax Simpson, 1928 [ Parendotherium Crusafont Pairó & Adrover, 1966 ]
  - View † L. valdensis (Woodward, 1911) Simpson, 1928 [ Dipriodon valdensis Woodward, 1911 ]
  - Species † L. herreroi (Crusafont Pairó & Adrover, 1966) [ Parendotherium herreroi Crusafont Pairó & Adrover 1966 ]
- Genus † Monobaatar Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
  - Species † M. mimicus Kielan-Jaworowska, Dashzeveg & Trofimov, 1987
- Genus † Sinobaatar Hu & Wang, 2002
  - Species † S. lingyuanensis Hu & Wang, 2002
  - Species † S. xiei Kusuhashi et al. , 2009
  - Species † S. fuxinensis Kusuhashi et al. , 2009
- Genus † Tedoribaatar Kusuhashi et al. , 2008
  - Species † T. reini Kusuhashi et al. , 2008
- Genus † Teutonodon Martin et al. , 2016
  - Species † Teutonodon langenbergensis Martin et al. 2016
Family † Albionbaataridae Kielan-Jaworowska & Ensom, 1994
- Genus † Albionbaatar Kielan-Jaworowska & Ensom, 1994
  - Species † A. denisae Kielan-Jaworowska & Ensom, 1994
- Genus † Kielanobaatar Kusuhashi et al. , 2010
  - Species † K. badaohaoensis Kusuhashi et al. , 2010
- Genus Proalbionbaatar Hahn & Hahn, 1998
  - Species † P. plagiocyrtus Hahn & Hahn, 1998
Suborder † Gondwanatheria McKenna 1971 [Gondwanatheroidea Krause & Bonaparte 1993 ]
- Family † Groeberiidae Patterson, 1952
  - Genus † Groeberia Patterson 1952
    - Species † G. minoprioi Ryan Patterson , 1952
    - View † G. pattersoni GG Simpson , 1970
  - Genus † Klohnia Flynn & Wyss 1999
    - View † K. charrieri Flynn & Wyss 1999
    - Species † K. major Goin et al. , 2010
  - Genus? † Epiklohnia Goin et al. , 2010
    - Species † Epiklohnia verticalis Goin et al. , 2010
  - Genus? † Praedens Goin et al. , 2010
    - Species † Praedens aberrans Goin et al. , 2010
- Family † Ferugliotheriidae Bonaparte, 1986
  - Genus † Ferugliotherium Bonaparte, 1986a [ Vucetichia Bonaparte, 1990 ]
    - † Ferugliotherium windhauseni Bonaparte, 1986a [ Vucetichia gracilis Bonaparte, 1990 ]
  - Genus † Trapalcotherium Rougier et al. , 2008
    - † Trapalcotherium matuastensis Rougier et al. , 2008
- Family † Sudamericidae Scillato-Yané & Pascual, 1984 [Gondwanatheridae Bonaparte, 1986 ; Patagonidae Pascual & Carlini, 1987 ]
  - Genus † Greniodon Goin et al. , 2012
    - † Greniodon sylvanicus Goin et al. , 2012
  - Genus † Vintana Krause et al. , 2014
    - † Vintana sertichi Krause et al. , 2014
  - Genus † Dakshina Wilson, Das Sarama & Anantharaman, 2007
    - † Dakshina jederi Wilson, Das Sarama & Anantharaman, 2007
  - Genus † Gondwanatherium Bonaparte, 1986
    - † Gondwanatherium patagonicum Bonaparte, 1986
  - Genus † Sudamerica Scillato-Yané & Pascual, 1984
    - † Sudamerica ameghinoi Scillato-Yané & Pascual, 1984
  - Genus † Lavanify Krause et al. 1997
    - † Lavanify miolaka Krause et al. 1997
  - Genus † Bharattherium Prasad et al. , 2007
    - † Bharattherium bonapartei Prasad et al. ,, 2007
  - Genus † Patagonia Pascual & Carlini '1987
    - † Patagonia peregrina Pascual & Carlini '1987
Suborder † Cimolodonta McKenna, 1975
- Genus? † Allocodon non Marsh, 1881
  - View † A. fortis Marsh, 1889
  - Species † A. lentus Marsh, 1892 [ Cimolomys lentus ]
  - Species † A. pumilis Marsh, 1892 [ Cimolomys pumilus ]
  - Species † A. rarus Marsh, 1889
- Genus? † Ameribaatar Eaton & Cifelli, 2001
  - Species † A. zofiae Eaton & Cifelli, 2001
- Genus? † Bubodens Wilson, 1987
  - View † Bubodens magnus Wilson, 1987
- Genus? † Clemensodon Krause, 1992
  - Species † Clemensodon megaloba Krause, 1992 [ Kimbetohia cambi , in partim]
- Genus? † Fractinus Higgins 2003
  - Species † Fractinus palmorum Higgins, 2003
- Genus? † Uzbekbaatar Kielan-Jaworowska & Nesov, 1992
  - View † Uzbekbaatar kizylkumensis Kielan-Jaworowska & Nesov, 1992
- Genus? † Viridomys Fox 1971
  - View † Viridomys orbatus Fox 1971
- Family † Corriebaataridae Rich et al. , 2009
  - Genus? † Corriebaatar Rich et al. , 2009
    - Species † Corriebaatar marywaltersae Rich et al. , 2009
- Paracimexomys group
  - Genus Paracimexomys Archibald, 1982
    - View? † P. crossi Cifelli, 1997
    - View? † P. dacicus Grigorescu & Hahn, 1989
    - View? † P. oardaensis (Codrea et al. , 2014) [ Barbatodon oardaensis Codrea et al. , 2014 ]
    - Species † P. magnus (Sahni, 1972) Archibald, 1982 [ Cimexomys magnus Sahni, 1972 ]
    - Species † P. magister (Fox, 1971) Archibald, 1982 [ Cimexomys magister Fox, 1971 ]
    - Species † P. perplexus Eaton & Cifelli, 2001
    - Species † P. robisoni Eaton & Nelson, 1991
    - Species † P. priscus (Lillegraven, 1969) Archibald, 1982 [ Cimexomys priscus Lillegraven, 1969 ; genotype Paracimexomys sensu Eaton & Cifelli, 2001 ]
    - Species † P. propriscus Hunter, Heinrich & Weishampel 2010
  - Genus Cimexomys Sloan & Van Valen, 1965
    - Species † C. antiquus Fox, 1971
    - Species † C. gregoryi Eaton, 1993
    - Species † C. judithae Sahni, 1972 [ Paracimexomys judithae (Sahni, 1972) Archibald, 1982 ]
    - Species † C. arapahoensis Middleton & Dewar, 2004
    - Species † C. minor Sloan & Van Valen, 1965
    - View? † C. gratus (Jepson, 1930) Lofgren, 1995 [ Cimexomys hausoi Archibald, 1983 ; Eucosmodon gratus Jepson, 1930 ; Mesodma ambigua ? Jepson, 1940 ; Stygimus gratus Jepson, 1930 ]
  - Genus † Bryceomys Eaton, 1995
    - Species † B. fumosus Eaton, 1995
    - Species † B. hadrosus Eaton, 1995
    - Species † B. intermedius Eaton & Cifelli, 2001
  - Genus † Cedaromys Eaton & Cifelli, 2001
    - Species † C. bestia (Eaton & Nelson, 1991) Eaton & Cifelli, 2001 [ Paracimexomys bestia Eaton & Nelson, 1991]
    - View † C. hutchisoni Eaton 2002
    - Species † C. minimus Eaton 2009
    - Species † C. parvus Eaton & Cifelli, 2001
  - Genus † Dakotamys Eaton, 1995
    - View? † D. sp. Eaton 1995
    - Species † D. malcolmi Eaton, 1995
    - View † D. shakespeari Eaton 2013
- Family † Boffidae Hahn & Hahn, 1983 sensu Kielan-Jaworowska & Hurum 2001
  - Genus † Boffius Vianey-Liaud, 1979
    - Species † Boffius splendidus Vianey-Liaud, 1979 [Boffiidae Hahn & Hahn, 1983 sensu Kielan-Jaworowska & Hurum, 2001 ]
- Family † Cimolomyidae Marsh, 1889 sensu Kielan-Jaworowska & Hurum, 2001
  - Genus † Paressodon Wilson, Dechense & Anderson, 2010
    - View † Paressodon nelsoni Wilson, Dechense & Anderson, 2010
  - Genus † Cimolomys Marsh, 1889 [? Allacodon Marsh, 1889 ; Selenacodon Marsh, 1889 ]
    - Species † C. clarki Sahni, 1972
    - Species † C. gracilis Marsh, 1889 [ Cimolomys digona Marsh, 1889 ; Meniscoessus brevis ; Ptilodus gracilis Osborn, 1893 non Gidley 1909 ; Selenacodon brevis Marsh, 1889 ]
    - Species † C. trochuus Lillegraven, 1969
    - Species † C. milliensis Eaton, 1993a
    - Species? † C. bellus Marsh, 1889
  - Genus? † Essonodon Simpson, 1927
    - Species † E. browni Simpson, 1927 [cimolodontidae? Kielan-Jaworowska & Hurum 2001 ]
  - Genus? † Buginbaatar Kielan-Jaworowska & Sochava, 1969
    - View † Buginbaatar transaltaiensis Kielan-Jaworowska & Sochava, 1969
  - Genus? † Meniscoessus Cope, 1882 [ Dipriodon Marsh, 1889 ; Tripriodon Marsh, 1889 nomen dubium ; Triprotodon Chure & McIntosh, 1989 nomen dubium ; Selenacodon Marsh, 1889 , Halodon Marsh, 1889 , Oracodon Marsh, 1889 ]
    - Species † M. caperatus Marsh, 1889
    - Species † M. collomensis Lillegraven, 1987
    - Species † M. conquistus Cope 1882
    - Species † M. ferox Fox, 1971a
    - Species † M. intermedius Fox, 1976b
    - Species † M. major (Russell, 1936) [ Cimolomys major Russell 1937 ]
    - Species † M. robustus (Marsh, 1889) [ Dipriodon robustus Marsh 1889 ; Dipriodon lacunatus Marsh, 1889 ; Tripriodon coelatus Marsh, 1889 ; Meniscoessus coelatus Marsh, 1889 ; Selenacodon fragilis Marsh, 1889 ; Meniscoessus fragilis Marsh, 1889 ; Halodon sculptus (Marsh, 1889) ; Cimolomys sculptus Marsh, 1889 ; Meniscoessus sculptus Marsh, 1889 ; Oracodon anceps Marsh, 1889 ; Oracodon conulus Marsh, 1892 ; Meniscoessus borealis Simpson, 1927c ; Meniscoessus greeni Wilson, 1987 ]
    - Species † M. seminoensis Eberle & Lillegraven, 1998a
- Family † Kogaionidae Rãdulescu & Samson, 1996
  - Genus † Kogaionon Rãdulescu & Samson, 1996
    - View † K. ungureanui Rãdulescu & Samson, 1996
  - Genus Hainina Vianey-Liaud, 1979
    - Species † H. belgica Vianey-Liaud, 1979
    - View † H. godfriauxi Vianey-Liaud, 1979
    - Species † H. pyrenaica Peláez-Campomanes, López-Martínez, Álvarez-Sierra & Daams, 2000
    - View † H. vianeyae Peláez-Campomanes, López-Martínez, Álvarez-Sierra & Daams, 2000
  - Genus † Barbatodon Rãdulescu & Samson, 1986
    - Species † B. transylvanicum Rãdulescu & Samson, 1986
- Family † Eucosmodontidae Jepsen, 1940 sensu Kielan-Jaworowska & Hurum, 2001 [Eucosmodontidae: Eucosmodontinae Jepsen, 1940 sensu McKenna & Bell, 1997 ]
  - Genus † Eucosmodon Matthew & Granger, 1921
    - Species † E. primus Granger & Simpson, 1929
    - Species † E. americanus Cope, 1885
    - Species † E. molestus Cope, 1869 [ Neoplagiaulax molestus Cope, 1869 ]
  - Genus Stygimys Sloan & Van Valen, 1965
    - View † S. camptorhiza Johnston & Fox, 1984
    - Species † S. cupressus Fox, 1981
    - Species † S. kuszmauli [ Eucosmodon kuszmauli ]
    - View † S. jepseni Simpson, 1935
    - View † S. teilhardi Granger & Simpson, 1929
- Family † Microcosmodontidae Holtzman & Wolberg, 1977 [Eucosmodontidae: Microcosmodontinae Holtzman & Wolberg, 1977 sensu McKenna & Bell, 1997 ]
  - Genus † Pentacosmodon Jepsen, 1940
    - Species † P. pronus Jepsen, 1940 [Djadochtatheroid? (Kielan-Jaworowska & Hurum, 2001) ]
  - Genus Acheronodon Archibald, 1982
    - Species † A. garbani Archibald, 1982
  - Genus † Microcosmodon Jepsen, 1930
    - Species † M. conus Jepsen, 1930
    - Species † M. rosei Krause, 1980
    - Species † M. arcuatus Johnston & Fox, 1984
    - Species † M. woodi Holtzman & Wolberg, 1977 [Eucosmodontine?]
    - Species † M. harleyi Weil, 1998
- Superfamily † Ptilodontoidea Cope, 1887 sensu McKenna & Bell, 1997 e Kielan-Jaworowska & Hurum, 2001
  - Family † Cimolodontidae Marsh, 1889 sensu Kielan-Jaworowska & Hurum, 2001
    - Genus † Liotomus Lemoine, 1882 [ Neoctenacodon Lemoine 1891 ]
      - View? † L. marshi (Lemoine, 1882) Cope, 1884 [ Neoctenacodon marshi Lemoine, 1882 ; Neoplagiaulax marshi (Lemoine 1882) ; Plagiaulax marshi (Lemoine 1882) ] [Eucosmodontidae? McKenna & Bell, 1997]
    - Genus † Yubaatar Xu et al. 2015
      - Species † Yubaatar zhongyuanensis Xu et al. 2015
    - Genus † Anconodon Jepsen, 1940
      - View? † A. lewisi (Simpson 1935) Sloan, 1987
      - Species † A. gibleyi ( Simpson, 1935 ) [ Ptilodus gidleyi Simpson, 1935 ]
      - Species † A. cochranensis ( Russell, 1929 ) [ Liotomus russelli (Simpson, 1935) ; Anconodon russelli (Simpson, 1935) Sloan, 1987 ; Ectopodon cochranensis (Russell, 1967) ]
    - Genus † Cimolodon Marsh, 1889 [ Nanomys Marsh, 1889 , Nanomyops Marsh, 1892]
      - Species † C. agilis Marsh, 1889
      - Species † C. foxi Eaton, 2002
      - Species † C. gracilis Marsh, 1889
      - Species † C. electus Fox, 1971
      - Species † C. nitidus Marsh, 1889 [ Allacodon rarus Marsh, 1892 sensu Clemens, 1964a ; Nanomys minutus Marsh, 1889 ; Nanomyops minutus (Marsh, 1889) Marsh, 1892 ; Halodon serratus Marsh, 1889 ; Ptilodus serratus (Marsh, 1889) Gidley 1909 ]
      - Species † C. parvus Marsh, 1889
      - Species † C. peregrinus Donohue, Wilson & Breithaupt, 2013
      - Species † C. similis Fox, 1971
      - Species † C. wardi Eaton, 2006
  - Family Incertae sedis
    - Genus Neoliotomus Jepsen, 1930
      - Species † N. conventus Jepsen, 1930
      - Species † N. ultimus ( Granger & Simpson, 1928 )
  - Family † Neoplagiaulacidae Ameghino, 1890 [Ptilodontidae: Neoplagiaulacinae Ameghino, 1890 sensu McKenna & Bell, 1997 ]
    - Genus † Mesodma Jepsen, 1940
      - View? † M. hensleighi Lillegraven, 1969
      - View? † M. senecta Fox, 1971
      - Species † M. ambigua Jepsen, 1940
      - View? † M. pygmaea Sloan, 1987
      - Species † M. formosa (Marsh, 1889) [Halodon formosus Marsh, 1889]
      - Species † M. primaeva (Lambe, 1902)
      - Species † M. thompsoni Clemens, 1964
    - Genus Ectypodus Matthew & Cranger, 1921 [Charlesmooria Kühne, 1969]
      - Species † E. aphronorus Sloan, 1981
      - View? † E. childei Kühne, 1969
      - View? † E. elaphus Scott, 2005
      - View? † E. lovei (Sloan, 1966) Krishtlaka & Black, 1975
      - Species † E. musculus Matthew & Granger, 1921
      - Species † E. powelli Jepsen, 1940
      - View? † E. simpsoni Jepsen, 1930
      - Species † E. szalayi Sloan, 1981
      - Species † E. tardus Jepsen, 1930
    - Genus † Mimetodon Jepsen, 1940
      - Species † M. krausei Sloan, 1981
      - Species † M. nanophus Holtzman, 1978 [ Neoplagiaulax nanophus Holtzman, 1978]
      - Species † M. siberlingi (Simpson, 1935) Schiebout, 1974
      - View † M. churchilli Jepsen, 1940
    - Genus † Neoplagiaulax Lemoine, 1882
      - Species † N. annae Vianey-Liaud, 1986
      - View? † N. burgessi Archibald, 1982
      - Species † N. cimolodontoides Scott, 2005
      - Species † N. copei Lemoine, 1885
      - Species † N. donaldorum Scott & Krause, 2006
      - Species † N. eocaenus Lemoine, 1880
      - View † N. grangeri Simpson, 1935
      - View † N. hazeni Jepsen, 1940
      - Species † N. hunteri Krishtalka, 1973
      - Species † N. jepi Sloan, 1987
      - Species † N. kremnus Johnston & Fox, 1984
      - Species † N. macintyrei Slaon, 1981
      - Species † N. macrotomeus Wilson, 1956
      - Species † N. mckennai Sloan, 1987
      - Species † N. nelsoni Sloan, 1987
      - Species † N. nicolai Vianey-Liaud, 1986
      - Species † N. paskapooensis Scott, 2005
      - View? † N. serrator Scott, 2005
      - View † N. sylvani Vianey-Liaud, 1986
    - Genus † Parectypodus Jepsen, 1930
      - View † P. armstrongi Johnston & Fox, 1984
      - View? † P. corystes Scott, 2003
      - View? † P. Foxi Storer, 1991
      - Species † P. laytoni Jepsen, 1940
      - Species † P. lunatus Krause, 1982 [ P. childei Kühne, 1969]
      - View † P. simpsoni Jepsen, 1940
      - Species † P. sinclairi Simpson, 1935
      - Species † P. sloani Schiebout, 1974
      - Species † P. trovessartianus Cope, 1882 [ P. trouessarti ; Ptilodus ; Mimetodon ; Neoplagiaulax ]
      - Species † P. sylviae Rigsby, 1980 [Ectypodus sylviae Rigby, 1980 ]
      - View? † P. vanvaleni Sloan, 1981
    - Genus † Cernaysia Vianey-Liaud, 1986
      - Species † C. manueli Vianey-Liaud, 1986
      - Species † C. davidi Vianey-Liaud, 1986
    - Genus † Krauseia Vianey-Liaud, 1986
      - Species † K. clemensi Sloan, 1981 [ Parectypodus clemensi Sloan, 1981 ]
    - Genus Xyronomys Rigby, 1980
      - Species † X. swainae Rigby, 1980 [Xironomys (sic); ? Eucosmodontidae]
    - Genus Xanclomys Rigby, 1980
      - View † X. mcgrewi Rigby, 1980
    - Genus † Mesodmops Tong & Wang, 1994
      - Species † M. dawsonae Tong & Wang, 1994
  - Family † Ptilodontidae Cope, 1887 [Ptilodontidae: Ptilodontinae Cope, 1887 sensu McKenna & Bell, 1997 ]
    - Genus † Kimbetohia Simpson, 1936
      - View † K. cambi [ Granger, Gregory & Colbert in Matthew, 1937, or Simpson, 1936 ]
      - View † K. sp. cf. K. cambi
    - Genus † Ptilodus Cope, 1881 [Chirox Cope, 1884 ]
      - View? † P. fractus
      - Species † P. kummae Krause, 1977
      - Species † P. gnomus Scott, Fox & Youzwyshyn, 2002 [cf. Ectypodus hazeni (Jepsen, 1940) Gazin, 1956 ]
      - Species † P. mediaevus Cope, 1881 [ Ptilodus plicatus (Cope, 1884) ; Chirox plicatus Cope, 1884 P. ferronensis Gazin, 1941 ]
      - Species † P. montanus Douglass, 1908 [ P. gracilis Gidley, 1909 ; P. admiralis Hay, 1930 ]
      - Species † P. tsosiensis Sloan, 1981
      - Species † P. wyomingensis Jepsen, 1940
    - Genus † Baiotomeus Krause, 1987
      - Species † B. douglassi Simpson, 1935 [ Ptilodus ; Mimetodon ; Neoplagiaulax ]
      - Species † B. lamberti Krause, 1987
      - Species † B. russelli Scott, Fox & Youzwyshyn, 2002
      - Species † B. rhothonion Scott, 2003
    - Genus Prochetodon Jepsen, 1940
      - Species † P. cavus Jespen, 1940
      - Species † P. foxi Krause, 1987
      - Species † P. taxus Krause, 1987
      - View? † P. speirsae Scott, 2004
- Superfamily † Taeniolabidoidea Granger & Simpson, 1929 sensu Kielan-Jaworowska & Hurum, 2001
  - Genus † Prionessus Matthew & Granger, 1925
    - Species † P. lucifer Matthew & Granger, 1925
  - † Lambdopsalidae Family
    - Genus † Lambdopsalis Chow & Qi, 1978
      - Species † L. bulla Chow & Qi, 1978
    - Genus: Sphenopsalis Matthew, Granger & Simpson, 1928
      - Species † S. nobilis Matthew, Granger & Simpson, 1928
  - † Taeniolabididae Granger & Simpson Family , 1929
    - Genus † Taeniolabis Cope, 1882
      - View † T. lamberti Simmons, 1987
      - Species † T. taoensis Cope, 1882
    - Genus † Kimbetopsalis
      - Species † K. simmonsae
- Superfamily † Djadochtatherioidea Kielan-Jaworowska & Hurum, 1997 sensu Kielan-Jaworowska & Hurum, 2001 [Djadochtatheria Kielan-Jaworowska & Hurum, 1997 ]
  - Kind? † Bulganbaatar Kielan-Jaworowska, 1974
    - View? † B. nemegtbaataroides Kielan-Jaworowska, 1974
  - Genus † Nemegtbaatar Kielan-Jaworowska, 1974
    - View? † N. gobiensis Kielan-Jaworowska, 1974
  - Family † Chulsanbaataridae Kielan-Jaworowska, 1974
    - Genus † Chulsanbaatar Kielan-Jaworowska, 1974
      - Species † C. vulgaris Kielan-Jaworowska, 1974
  - Family † Sloanbaataridae Kielan-Jaworowska, 1974
    - Genus † Kamptobaatar Kielan-Jaworowska, 1970
      - View? † K. kuczynskii Kielan-Jaworowska, 1970
    - Genus † Nessovbaatar Kielan-Jaworowska & Hurum, 1997
      - Species † N. multicostatus Kielan-Jaworowska & Hurum, 1997
    - Genus † Sloanbaatar Kielan-Jaworowska, 1974
      - Species † S. mirabilis Kielan-Jaworowska, 1974 [Sloanbaatarinae]
  - Family † Djadochtatheriidae Kielan-Jaworowska $ Hurum, 1997
    - Genus † Djadochtatherium Simpson, 1925
      - View † D. matthewi Simpson, 1925 [ Catopsalis matthewi Simpson, 1925 ]
    - Genus † Catopsbaatar Kielan-Jaworowska, 1974
      - Species † C. catopsaloides (Kielan-Jaworowska, 1974) Kielan-Jaworowska, 1994 [ Djadochtatherium catopsaloides Kielan-Jaworowska, 1974 ]
    - Genus † Tombaatar Kielan-Jaworowska, 1974
      - Species † T. sabuli Rougier, Novacek & Dashzeveg, 1997
    - Genus † Kryptobaatar Kielan-Jaworowska, 1970 [ Gobibaatar Kielan-Jaworowska, 1970 , Tugrigbaatar Kielan-Jaworowska & Dashzeveg, 1978 ]
      - View † K. saichanensis Kielan-Jaworowska & Dashzeveg, 1978 [ Tugrigbaatar saichaenensis Kielan-Jaworowska & Dashzeveg, 1978 ?? ]
      - View † K. dashzevegi Kielan-Jaworowska, 1970
      - Species † K. mandahuensis Smith, Guo & Sun, 2001
      - View † K. gobiensis Kielan-Jaworowska, 1970 [ Gobibaatar parvus Kielan-Jaworowska, 1970 ]

Alternative Classification

The cladogram of subordinate taxa up to and including families is given according to the Paleobiology Database website for April 2018 ^[2] :

Allotheria

Euharamiyida

Gondwanatheria

Multituberculata

Arginbaataridae Hahn & Hahn, 1983

Hahnotheriidae Butler & Hooker, 2005

Haramyidae

Kermackodontidae Butler & Hooker, 2005

Cimolodonta McKenna, 1975

Boffiidae Hahn & Hahn, 1983

Cimolomyidae Sloan & Van Valen, 1965

Corriebaataridae Rich et al. , 2009

Eucosmodontidae Jepsen, 1940

Kogaionidae Rãdulescu & Samson, 1996

Microcosmodontidae Holtzman & Wolberg, 1977

Neoplagiaulacidae Ameghino, 1890

Djadochtatherioidea Kielan-Jaworowska & Hurum, 2001

	Djadochtatheriidae Kielan-Jaworowska & Hurum, 1997

	Sloanbaataridae Kielan-Jaworowska, 1974

Ptilodontoidea Sloan & Van Valen, 1965

	Cimolodontidae Marsh, 1889

	Ptilodontidae Cope, 1887

Taeniolabidoidea McKenna, 1975

	Lambdopsalidae Chow & Qi, 1978

	Taeniolabididae Granger & Simpson, 1929

Plagiaulacida Ameghino, 1889

	Albionbaataridae Kielan-Jaworowska & Ensom, 1994

	Allodontidae Marsh, 1889

	Eobaataridae Kielan-Jaworowska, Dashzeveg & Trofimov, 1987

	Paulchoffatiidae Hahn, 1969

	Pinheirodontidae Hahn & Hahn, 1999

	Plagiaulacidae Gill, 1872

	Zofiabaataridae Bakker, 1992

Notes

↑ Multi-tuberous / Lopatin A.V. // Meot archaeological culture - Mongol-Tatar invasion. - M .: Big Russian Encyclopedia, 2012. - S. 536-537. - (The Big Russian Encyclopedia : [in 35 vols.] / Ch. Ed. Yu. S. Osipov ; 2004—2017, vol. 20). - ISBN 978-5-85270-354-5 .
↑ ¹ ² Multituberculata (English) information on the Paleobiology Database website. (Retrieved May 27, 2018) .
↑ Weil 1997
↑ Meng Chen, Gregory Philip Wilson, A multivariate approach to infer locomotor modes in Mesozoic mammals, Article in Paleobiology 41 (02) February 2015 DOI: 10.1017 / pab.2014.14
↑ Agustí-Antón 2002, pp. 3-4.
↑ Benton, Michael J. Vertebrate Palaeontology (2004), p. 300 .
↑ Carrano, Matthew T., and Richard W. Blob, Timothy J. Gaudin, and John R. Wible (2006). Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles , p. 358 .
↑ Mikko's Phylogeny Archive [1] Haaramo, Mikko Mammaliaformes - mammals and near-mammals (neopr.) (2007). Date of treatment December 30, 2015.
↑ Paleofile.com (net, info) [2] . Taxonomic lists- Mammals (neopr.) . Date of treatment December 30, 2015.

Literature

Butler PM & Hooker JJ , 2005: New teeth of allotherian mammals from the English Bathonian, including the earliest multituberculates. Acta Palaeontologica Polonica: Vol. 50. Documento pdf
Hahn G. & Hahn R. , 2003: New multituberculate teeth from the Early Cretaceous of Marocco. Acta Palaeontologica Polonica: Vol. 48.
Kielan-Jaworowska Z. , Cifelli RL & Luo Z.-X. 2004: Mammals from the Age of Dinosaurs - origins, evolution, and structure. Columbia University Press, New York, 2004.
McKenna MC & Bell SK , (eds.) 1997: Classification of mammals - above the species level. Columbia University Press, New York, 1997.
Kielan-Jaworowska Z. and Hurum JH (2001), "Phylogeny and Systematics of multituberculate mammals". Paleontology 44 , p. 389-429.

Links

Introduction .

[BRE-1] Multi-tuberous / Lopatin A.V. // Meot archaeological culture - Mongol-Tatar invasion. - M .: Big Russian Encyclopedia, 2012. - S. 536-537. - (The Big Russian Encyclopedia : [in 35 vols.] / Ch. Ed. Yu. S. Osipov ; 2004—2017, vol. 20). - ISBN 978-5-85270-354-5 .

[PBDB-2] ¹ ² Multituberculata (English) information on the Paleobiology Database website. (Retrieved May 27, 2018) .

[Weil-3] Weil 1997

[4] Meng Chen, Gregory Philip Wilson, A multivariate approach to infer locomotor modes in Mesozoic mammals, Article in Paleobiology 41 (02) February 2015 DOI: 10.1017 / pab.2014.14

[Agusti-3-5] Agustí-Antón 2002, pp. 3-4.

[6] Benton, Michael J. Vertebrate Palaeontology (2004), p. 300 .

[7] Carrano, Matthew T., and Richard W. Blob, Timothy J. Gaudin, and John R. Wible (2006). Amniote Paleobiology: Perspectives on the Evolution of Mammals, Birds, and Reptiles , p. 358 .

[mikko-8] Mikko's Phylogeny Archive [1] Haaramo, Mikko Mammaliaformes - mammals and near-mammals (neopr.) (2007). Date of treatment December 30, 2015.

[9] Paleofile.com (net, info) [2] . Taxonomic lists- Mammals (neopr.) . Date of treatment December 30, 2015.